Chemical physiology, pheromone communication, fish pheromone regulation, fish reproduction and stress endocrine regulation, jawless fish evolution, genome annotation and functional genomics, invasive species control. He has successively presided over the fund support of NIH, NSF, US Department of Energy, SEA GRANT, Great Lakes Fisheries Commission, Great Lakes Fisheries Trust Fund, Great Lakes Conservation Fund, and Canadian Science and Engineering Research Department (NSERC), with a total fund of over US$ 654.38+065.438+000,000. In recent years, many papers with strong academic influence have been published in top international academic journals such as Science, PNAS, Journal of Neuroscience and BMC Evolutionary Biology.